En Sabah Nur is an ancient and immortal mutant with an array of superhuman abilities ranging from shapeshifting to immense superhuman strength and durability. Apocalypse's ultimate goal is to eradicate all life from Earth in order to reshape it in his image. He currently serves in the Autumn section of the Quiet Council of Krakoa and is an ally to Excalibur.
Dominikos Petrakis is a mutant with the ability to generate seismic waves from his hands. He most often uses this ability to create tremors and earthquakes. Avalanche was first recruited into Mystique's Brotherhood of Evil Mutants and has remained a prominent X-Men villain since.
A second, unnamed Avalanche appeared after the apparent death of Petrakis.
Thomas Cassidy is a mutant with the ability to channel bio-kinetic energy into concussive blasts with wooden objects and later gained the ability to manipulate plant life. Black Tom is often seen with his partner, Juggernaut. He is currently a member of Krakoa's mutant intelligence group X-Force.
Frederick J. Dukes is a mutant with elastic, blubbery skin. Morbidly obese, Blob is immune to most physical harm. He also possesses a personal gravity field which enabled him to render himself immovable. Blob was a member of the original Brotherhood of Evil Mutants and has been one of the X-Men's most recurring and persistent adversaries.
Although Blob appeared to lose his powers after Decimation, he has since appeared with his powers intact.
Cal'Syee Neramani is a Shi'ar warrior and Empress Lilandra's older sister. Exiled from the Shi'ar Empire, Deathbird sought to reclaim her throne which put her at odds with Lilandra's allies, the X-Men.
Bennet du Paris is a powerful mutant with telepathy, telekinesis, teleportation, and immortality. Exodus was a loyal member of the Acolytes, worshipers of Magneto, for many years. He currently serves the Winter section of the Quiet Council of Krakoa.
Cain Marko is Professor Charles Xavier's stepbrother and one of the X-Men's earliest enemies. After finding the mystical gem of Cyttorak, Marko is empowered and becomes the Juggernaut, a massive, unstoppable creature capable of shattering mountains. Despite lacking the mutant gene, Juggernaut is often a member of various incarnations of the Brotherhood of Mutants.
Yuriko Oyama is the daughter of Japanese crime boss Lord Dark Wind. After the death of her lover, Oyama sought out Spiral in order to enhance her body. She was modified with adamantium bones and talons, and took the name Lady Deathstrike.
Max "Erik Magnus" Eisenhardt is a mutant with the ability to generate and control magnetic fields. A close friend of Professor X, Magneto clashed with the X-Men over the philosophy that mutants and humans could coexist. A survivor of The Holocaust, Magneto believed mutants were superior to humans. Magneto has formed several groups over the years to fight for mutant supremacy, including the Brotherhood of Mutants and the Acolytes. Magneto has also served with the X-Men on several occasions and is currently one of the central decision-makers on the mutant nation of Krakoa, serving in the Autumn section of the Quiet Council of Krakoa.
Jason Wyngarde was a mutant with the power to create illusions. A member of several villain groups such as the Brotherhood of Mutants, Factor Three, and the Hellfire Club, Mastermind is largely known for being responsible for Jean Grey's transition into the Dark Phoenix. He later died from the Legacy Virus after he begs for Jean Grey's forgiveness. Mastermind was later resurrected when the X-Men relocated to Krakoa.
Nathaniel Essex is a genetically-altered human and scientific genius, and one of the X-Men's deadliest enemies. Sinister first rose to prominence when he ordered his assassins, the Marauders, to hunt down and kill the sewer-dwelling mutants known as the Morlocks. He later played key roles as a mastermind in the shadows through events such as Inferno and Messiah CompleX. Mister Sinister has mastered genetic cloning, which he uses to create clones of the Marauders and others.
Although not a mutant at birth, Essex genetically engineered the X-Gene of deceased X-Man John Proudstar into his own genome. In exchange for his collection of mutant DNA he's collected over the years, he was welcomed to the mutant nation of Krakoa, where he serves in the Winter section of the Quiet Council of Krakoa. Outside of his duties in the council, he is also a field leader of sorts of the Hellions.
An obese, yellow-skinned alien from a race known as the “Spineless Ones,” Mojo is a slaver who rules the Mojoverse, a dimension obsessed with television programs and celebrities.
Raven Darkholme is a mutant with shapeshifting abilities. While she first appeared as a Ms. Marvel villain, Mystique has become one of the most iconic and recurring adversaries for the X-Men. She currently serves in the Winter section of the Quiet Council of Krakoa.
St. John Allerdyce is an Australian mutant with pyrokinesis, the ability to manipulate flame. Pyro has served with multiple incarnations of the Brotherhood of Mutants, as well as the government-led Freedom Force. Pyro later contracted the Legacy Virus and died months after. However, Pyro was resurrected by Selene during Necrosha, and has made multiple appearances since. He later died once again, but was the first mutant resurrected by the X-Men as they took residence on the mutant island of Krakoa. There, Pyro joined Kate Pryde's band of mutant pirates known as the Marauders.
A second Pyro named Simon Lasker joined Mesmero's Brotherhood of Mutants, but later reformed and joined the X-Men.
Victor Creed is a mutant with a regenerative healing factor similar to Wolverine's. Like Wolverine, Sabretooth gained an adamantium skeleton and claws under unknown circumstances. He was the first mutant to break the laws of the newly founded mutant nation Krakoa and was banished to a void within the island where he will live the rest of his life in permanent stasis.
Dr. Karl Lykos is a human geneticist who was transformed into a pterodactyl-like humanoid. As the winged Sauron, Lykos is an energy vampire who must absorb the lifeforce of those around him for sustenance. If Lykos absorbs a mutant's energy, he is transformed into Sauron, and his savage mind takes over.
Sebastian Shaw was a mutant with the ability to absorb kinetic and thermal energy directed at him. He was the leader of the New York branch of the Hellfire Club, where he has ruthlessly sacrificed mutant interests for the sake of money and power. He is currently the Black King of the Hellfire Trading Company and has employed his recently resurrected son Shinobi Shaw and the twin mercenaries Fenris as his Black Bishop and Black Knights, respectively, to grapple power of Krakoan trade from Emma Frost and Kate Pryde.
Amahl Farouk is an Egyptian mutant with powerful telepathic abilities and the power to possess other beings while on the Astral Plane. The Shadow King has played a key role in Storm's origin, and has often faced off against Professor X.
Rita Wayword is a human who was forcibly modified into the sorceress known as Spiral by her former master, Mojo. As Spiral, Wayword has powerful mystical abilities able to defeat the X-Men singlehandedly. She briefly served with Freedom Force and later opened a “Body Shoppe” where she modified others, such as Lady Deathstrike and the other Reavers, as well as the X-Men's Psylocke. On occasions, Mojo would summon Spiral to serve him, though she later retrieved the part of her soul stolen from her by Mojo which stripped Mojo's control over her and activated the complete scope of her powers.
Mortimer Toynbee is a mutant with superhuman leg and tongue strength and the ability to secrete a paralyzing venom from his mucus. Toad served as Magneto's henchman for several years until he formed his own version of the Brotherhood of Mutants. His leadership was temporary, however, as he resumed his status as an underling in later incarnations of the team.
Emma Frost is a mutant with powerful telepathic abilities, and later a secondary mutation that allows Frost to transform into an indestructible diamond form. While Emma Frost first appeared as the White Queen of the Hellfire Club, she later joined the X-Men and became a prominent member of the group.
Dr. Roderick “Rory” Campbell is a time-traveling cyborg from the future with the ability to generate energy harpoons. Ahab was a key figure in the origin of Rachel Summers, the future daughter of Cyclops of Jean Grey. He later served Apocalypse as his Horseman of Famine.
Bella Donna Boudreaux is a mutant with astral projection abilities, as well as the power to generate plasma blasts. A member of the Assassin's Guild, Bella Donna was a prominent figure in Gambit's past.
Bolivar Trask was an anthropologist who saw the rise of mutants as a threat to humanity. Trask was the developer of the robot guardians known as the Sentinels. Trask later realized the errors of his ways and sacrificed himself to destroy the Sentinels' base.
An albino mutant and former member of the sewer-dwelling Morlocks, Caliban had the ability to sense and track any mutant. Originally an ally to the X-Men, Caliban was corrupted by Apocalypse and transformed into a Horseman of Apocalypse shortly after the morlock massacre. He later overcame this corruption and joined the X-Men.
Leader of the underground-dwelling mutant community known as the Morlocks, Callisto is a mutant with enhanced senses. She is currently the White Knight of the Hellfire Trading Company, under the leadership of Emma Frost.
Cameron Hodge was a human who deceived X-Factor by posing as an ally. But in reality, he despised mutants and plotted to kill the members of X-Factor. He was later killed and bonded with the Phalanx.
Candra
Gambit #1 (December 1993)
A member of the Externals, Candra is a mutant with powerful telekinetic abilities and immortality. Candra is most often associated with the Assassins and Thieves Guilds where she is known as the Benefactress, bestowing great powers to those who serve her. She also has a history with Storm, who had stolen the gem known as the Heart of Candra, the source of her powers.
Ralph Roberts was a scientist studying cobalt and nuclear radiation. He used his research to make a suit similar to Iron Man planning to give it the military. When he first put it on, he was suffering from a concussion due to a pole jump injured, causing him to become obsessed with defeating Iron Man. He was defeated by the X-Men before his unstable suit could explode.
Frank Bohannan was a mutant with peak human physical prowess. Along with Super Sabre and Stonewall, Crimson Commando was a World War II veteran who joined Freedom Force. During a mission with Freedom Force, Crimson Commando was gravely injured and forced to undergo cybernetic modifications. He then changed his codename to simply Commando. After Decimation, he was one of the many mutants who lost their powers. He was later killed by Wolverine after kidnapping Hope Summers.
Silas Burr was a mutant with superhuman strength. He later modified his body with adamantium, granting himself indestructible skin. After Wolverine lost his adamantium, Genesis captured and killed Cyber in attempt to bond his adamantium with the X-Man's. Cyber later escaped Hell and became the new Hornet.
Danger
Astonishing X-Men (vol. 3) #9 (2005)
Danger is a sentient AI robot that is the physical manifestation of the Danger Room. It is agender, but has taken the form and shape of a female humanoid android.
Henry Phillip McCoy is a twisted version of the X-Man, Beast, from the Age of Apocalypse reality. Dark Beast escaped the Age of Apocalypse reality into Earth-616 and has since appeared as a major nemesis of the X-Men.
Wade Wilson is a mutated human (not a mutant) with regenerative abilities and enhanced strength. After facing terminal cancer, Wilson joined Canada's Weapon X program where he became the "merc with a mouth" known as Deadpool. Deadpool battled X-Force on several occasions, but later joined the X-Men and several variations of the team.
Irene Adler was a blind mutant with precognitive powers. Mystique's lover, Destiny foresaw many future events which she kept in a diary. Destiny was later killed in battle, though Mystique used trickery to have her revived using the Krakoan resurrection process.[1]
Donald Pierce is a cyborg and member of the Hellfire Club and the Reavers. Pierce joined the Hellfire Club to kill the other members of the Inner Circle, all of whom were mutants. He later created the cyborg mercenary group called the Reavers to battle the X-Men. He has since appeared as a member of both teams, alternating his allegiance to suit his whims.
Fabian Cortez is one of the founding members of the Acolytes, mutants devoted to Magneto. Cortez has the mutant ability to empower other mutants around him, though secretly he would slowly absorb his victims' lifeforces. Despite his many betrayals, Fabian Cortez was invited to the mutant island of Krakoa.
Joanna Cargill is a mutant with superhuman strength and indestructible skin. Frenzy has worked for the Alliance of Evil, the Femizons, the Acolytes, and the Marauders, though she recently reformed and joined the X-Men. She currently serves as Earth's ambassador and a member of S.W.O.R.D.
A mysterious mutant with powerful telepathic abilities. Alongside Selene, the Gamesmaster organized and created the Upstarts, a competition for wealthy young individuals to kill mutants.
Leader of band of the extraterrestrial bounty hunters known as Technet, Gatecrasher is a massive, blue-skinned female alien with telepathic abilities. She is most often associated as an antagonist for Captain Britain and Excalibur.
Gauntlet was the original leader of the Dark Riders. A member of the Inhumans, Gauntlet has been a member of each incarnation of the Dark Riders, though he was recently killed by Magneto.
Tyler Dayspring was the son of Cable who had previously taken the codename Tolliver. As Genesis, Dayspring took over leadership of the Dark Riders and plotted against Wolverine.
A member of the Externals and prominent adversary of X-Force. Gideon has the mutant ability to take on the powers of nearby mutants. He was killed by Selene along with the majority of other Externals, though he has since returned without explanation.
Radha Dastoor was a woman who believed that mutants and humans would become unified after a series of catastrophes. Haven possessed transdimensional powers that enabled her to teleport herself and others to her own pocket dimension, and battled X-Factor multiple times. She was later killed after giving birth to the Adversary.
Hazard
X-Men (vol. 2) #12 (September 1992)
Carter Ryking was a mutant with plasma generation. A childhood friend of Professor X, Ryking was experimented on by Mister Sinister at a young age. He later lost his powers during Decimation, which drove him insane. He was killed while in Ryking Institute, a mental asylum.
Holocaust
X-Men Alpha (February 1995)
Created by Scott Lobdell and Roger Cruz. Originally known as Nemesis,[2] Holocaust claimed to be the son of Apocalypse. In 2018, CBR.com ranked Holocaust 22nd in their "Age Of Apocalypse: The 30 Strongest Characters In Marvel's Coolest Alternate World" list,[3] and MSN included the character in the 2024 list of the "10 Strongest X-Men Villains" and rated him as "horrifyingly powerful mutant".[4] Scholar of religion and philosophy Gareth Evans-Jones observed that topics of persecution and injustice in the X-Men comics invite comparisons to historical events, so much so that "the concept of the 'holocaust' was later personified in [...] with the character Holocaust".[5]
Lani Ubanu is a mutate from the Savage Land with a hypnotic voice that can mesmerize male humanoids. While normally associated with the Savage Land, Lorelei is notable for being a member of one of Magneto's original Brotherhood of Evil Mutants lineups.
Calvin Montgomery Rankin is one of the X-Men's earliest members, and the first character to be added to the X-Men's original roster. Mimic has the mutant ability to copy the powers of those within his radius.
Lois London is a mutant and sister to the X-Men's Dazzler. During Necrosha, Mortis joined Selene's Inner Circle and helped her resurrect several deceased mutants.
Nanny
X-Factor #30 (July 1988)
Nanny was once a scientist employed by the Right in the field of cyborg technology. At some point, Nanny was captured by the X-Men, though her partner, Orphan-Maker, freed her. She then joined the Hellions on Krakoa.
A demon from the realm of Limbo. A former servant of Belasco, N'astirh coveted power and wished to become ruler of Limbo as well as Earth. Along with S'ym, N'astirh was responsible for the demonic event known as Inferno. He was later killed by Storm, though Magik has since resurrected the demon.
Eileen Harsaw is a mutant with the ability to disrupt the electromagnetic field of those around her. Toad recruited Phantazia into his Brotherhood of Evil Mutants, realizing how powerful she was. She was later invited to join the Acolytes, but she rejected the offer. Phantazia lost her powers after Decimation, and was last seen in a mental institution repeating the words “House of M”.
Kevin MacTaggert is a reality-warping and body-possessing mutant and the son of X-Men ally Moira MacTaggert. He is a member of The Five, a group of mutants whose abilities in tandem resurrect fallen mutants within the island of Krakoa.
A prominent United States Senator who began his career on an anti-mutant platform and tended to be an antagonist to the X-Men team. But towards the later days of his career, he began to change his views on mutants as a whole.
Opal Luna Saturnyne is an Omniversal Majestrix. She is cunning, deceitful and manipulative and she oversaw the battle in X of Swords between the mutants of Krakoa and mutants of Arakko.
Shinobi Shaw was a mutant with the power to alter his body's density from diamond-hard to intangible. The son of Sebastian Shaw of the Hellfire Club, Shinobi killed his father and took over his role as Black King while also working with the Upstarts, a group of individuals competing to score points by killing mutants.
After a number of deaths, Shinobi was resurrected by the X-Men on the mutant island Krakoa and recruited into the Hellfire Trading Company by his father.
Siena Blaze was a mutant with the ability to fire destructive blasts of electromagnetic energy. When she used her powers, she risked destroying the Earth. During her first outing, Siena Blaze nearly killed the three leaders of the X-Men in Antarctica. She was later killed at the Weapon X Neverland concentration camp, though she was resurrected by Selene much later.
A clone of Cable and major adversary of X-Force. From the same future timeline as Cable, Stryfe traveled to the past and formed the Mutant Liberation Front, a terrorist group. He was also primarily responsible for the release of the Legacy Virus, a fatal disease for mutantkind.
Trevor Fitzroy was a mutant with the ability to absorb the lifeforce of those he touches with his hands. For many years, Fitzroy served with the Upstarts. He was later killed by the Office of National Emergency.
Gunther Bain is a mutant with the ability to generate an invisible psionic field around himself. One of the X-Men's earliest foes, Unus joined a variation of Magneto's original Brotherhood of Mutants. Over the years, Unus was killed in action and later returned, but lost his powers after Decimation. Despite this, he recently appeared with his powers intact.
Telford Porter was a mutant with the power of teleportation. One of the X-Men's earliest foes, Vanisher had been a member of the Brotherhood of Evil Mutants, Factor Three, Marauders, and Hellfire Club. He later joined X-Force. Vanisher was shot and killed while facing off against General Callahan.
A superhuman being who considered himself to be “homo superior superior.” Vargas was killed by Harpoon of the Marauders while in possession of a piece of Destiny's diaries.
A mutate from the Savage Land with the ability to disrupt the equilibrium of living creatures. A clone of Vertigo was later recruited into the Marauders by Mister Sinister.
Gabriel Summers is a mutant with the ability to psionically manipulate, control, and absorb vast amounts of energy. Vulcan is the younger brother of X-Men members Cyclops and Havok.
Zaladane is a sorceress and possible mutate, most often associated with the Savage Land. While initially a priestess of Garokk, Zaladane later assisted the High Evolutionary and then afterward assumed the role of Empress of the Savage Land. Zaladane was later killed by Magneto against the protests of Rogue and Nick Fury.
A group of mutantfanatics who believe that mutants are destined to conquer and destroy normal humanity. The Acolytes were loyal and completely devoted to Magneto, who they believed to be the mutant messiah. The Acolytes were major X-Men foes throughout the 1990s, though the group has been seldom seen since.
A band of mutant mercenaries who have been hired by Apocalypse, A.I.M., and other criminals. The group was short-lived as Tower was killed by the X-Cutioner and Frenzy joined the Femizons and Acolytes, while Apocalypse abandoned the group for the Four Horsemen of Apocalypse.
A team of mutants founded by Magneto to serve as the antithesis to the X-Men. The team has had many incarnations over the years, with Mystique, Toad, Exodus, Mesmero, and others leading their own versions.
Other prominent members include Mastermind, Blob, Pyro, Avalanche, Destiny, Juggernaut, Phantazia, Vanisher, and Sauron.
The descendants of Apocalypse. They first appeared in Apocalypse vs Dracula #1. While Blink and Chamber are descendants of Apocalypse, neither is considered a member of the Clan.
Dark Descendants
The Uncanny X-Men #335 (August 1996)
A group of mutant worshipers of Onslaught led by Dark Beast.
Also known as the Riders of the Storm, the Dark Riders are servants of Apocalypse devoted to the philosophy of the survival of the fittest. The first incarnation was composed primarily of Inhumans, though when Genesis took over as leader of the second incarnation, many mutants were added to the roster.
A group of a rare, immortal subspecies of mutants. Prominent members include Candra, Gideon, Absalom, Crule, and Selene. Apocalypse was later revealed to be an External.
A group of four individuals (typically mutants) that have been genetically altered and mentally conditioned to serve the immortal mutant Apocalypse. The first modern incarnation included Warren Worthington III as Death, Autumn Rolfson as Famine, Abraham Kieros as War, and Plague as Pestilence. However, Apocalypse has had several incarnations of the Horsemen over the years.
A team of federal agents which was established as a superhero team directly accountable to the US government. The team was formed from former members of the Brotherhood of Evil Mutants's second incarnation, but almost immediately began taking on new recruits, including Spiral, Spider-Woman, Stonewall, Crimson Commando, and Super Sabre.
A group of female mercenaries led by Cyber. The Hell's Belles functioned as extortionists for high-paying drug cartels. Their membership included fire-breathing Flambé, scorching hot Briquette, invisible Vague, and the sonic wave-projecting Tremolo. The second incarnation of X-Factor was hired to protect former Belle Shrew from the group's attempts at revenge.
All of the Belles except for Briquette were depowered after Decimation. The depowered Belles were eventually invited to join Krakoa and potentially have their powers restored through the Crucible.
Due to the long-standing and secretive nature of the organization, not all of its members are known. The listed members are all part of the Inner Circle, also known as the Lords Cardinal. The non-Inner Circle members include many wealthy or politically connected men and women of the Marvel Universe like Tony Stark and Norman Osborn and are generally unaware and/or uninvolved with the Club's illegal activities.
Prominent members include Emma Frost, Sebastian Shaw, Donald Pierce, Harry Leland, Selene, Shinobi Shaw, Friedrich Von Roehm and Sage (then known as Tessa).
An anti-mutant organization focused on harvesting the X-Gene. Its prominent members are its director Lydia Nance and Edwin Martynec. The organization is a subsidiary of Orchis
Homines Verendi
Marauders #4 (December 2019)
Anti-mutant organization led by the children who once commandeered the Hellfire Club: Kade Kilgore, Manuel Enduque, Wilhelmina Kensington and Maximilian Frankenstein.
Hordeculture
X-Men (vol. 5) #3 (February 2020)
A group of elderly human scientists "who specialize in the genetic manipulation of – and propagation of – all things botanical." Membership includes Augusta Bromes, Edith Scutch, Opal Vetiver, and Lily Leymus. Former member includes Harrower.
A group of mutant mercenaries recruited by Mister Sinister to serve as his strike force. Each Marauder's DNA has been stored by Sinister so that in the event of their deaths, he may clone them at a moment's notice. Each Marauder retains the memories of the clone who died before them.
An underground society of mutant outcasts living in the sewers and abandoned tunnels beneath New York City, composed of mutant misfits, especially those with physical mutations or other conspicuous manifestations of their mutant genetics who were unable to pass as human in everyday society.
Orchis is a secretive anti-mutant organization composed of former members of A.I.M., Hydra, S.H.I.E.L.D., A.R.M.O.R., and Alpha Flight. Orchis created Mother Mold, an advanced version of Master Mold. When a team of X-Men attacked the Orchis Forge space station, they destroyed Mother Mold but perished in the resulting destruction. Orchis was also responsible for the events of "Fall of X" and has run afoul of other superheroes like Iron Man.
A team of cyborgs led by Lady Deathstrike and Donald Pierce. The Reavers were originally created by Spiral, transforming the mortally wounded Hellfire Club mercenaries Reese, Cole, and Macon into cyborgs to save their lives. When Donald Pierce joined the team, he brought with him Skullbuster, Pretty Boy, and Bonebreaker. Cylla Markham later joined the group to replace Skullbuster after his death.
Rising Sons
Generation X #53 (May 1999)
A team of adolescent Madripoor-based mercenaries composed of mutants, cyborgs and magic-users led by Dragonwing and including his sister, Spoilsport, along with The Sign, Jet-Black, Tough Love and Nightwind.
Primitive humans genetically altered by Magneto. When Magneto abandoned the Savage Land, the mutates were led by Sauron. The Sun Priestess Zaladane later took over as Empress of the Savage Land until her demise. The mutates eventually turned on Sauron, who had returned to lead, instead opting for a shared leadership between Lupa, Vertigo, and Whiteout.
Technology-advanced mutant-hunting robots created by Bolivar Trask
Swordbearers of Arakko
X-Men (vol. 5) #12 (November 2020)
A group of mutant inhabitants of Arakko, Krakoa's sister entity, who have each been given a mythical sword. Members include Summoner, Isca the Unbeaten, Bei the Blood Moon, Redroot the Forest, Solem, the White Sword, Pogg Ur-Pogg, and Apocalypse's first Horsemen of the Apocalypse.
A second version of Weapon P.R.I.M.E. later appeared, this time led by Tigerstryke. Weapon P.R.I.M.E. briefly battled X-Force and later attempted to hunt down Northstar.
XENO
X-Force (vol. 6) #1 (November 2019)
XENO is an extreme anti mutant organization based out of Seoul, South Korea. Its leader is the Man with the Peacock Tattoo.