^Segura MJ, Meyer MM, Matsuda SP (July 2000). "Arabidopsis thaliana LUP1 converts oxidosqualene to multiple triterpene alcohols and a triterpene diol". Organic Letters. 2 (15): 2257–9. doi:10.1021/ol006016b. PMID10930257.
^Kushiro T, Hoshino M, Tsutsumi T, Kawai K, Shiro M, Shibuya M, Ebizuka Y (November 2006). "Stereochemical course in water addition during LUP1-catalyzed triterpene cyclization". Organic Letters. 8 (24): 5589–92. doi:10.1021/ol062310d. PMID17107079.