Subsequent molecular phylogenetic studies upheld the relationship between the Theraphosidae and Barychelidae, but found that Paratropidae fell outside the clade.[2][3] In 2012, it was suggested that Theraphosoidina should probably include an additional family of spiders related to Homostola.[4] A major 2020 study of the Mygalomorphae established the new family Bemmeridae (raised from the rank of tribe) containing the two genera Homostola and Spiroctenus and included it within Theraphosoidina.[5]
Phylogeny
An outline version of the preferred cladogram from a 2020 phylogenetic study of the Mygalomorphae is shown below. (A node with lower support is marked ♦.) Although the precise position of the Theraphosoidina clade varied in some of the individual analyses, the clade itself was well supported.[5]
According to Opatova et al. (2020), the clade includes the following families, one of which (Bemmeridae) was first described at this rank in their study:[5]
^ abHedin, Marshal & Bond, Jason E. (2006), "Molecular phylogenetics of the spider infraorder Mygalomorphae using nuclear rRNA genes (18S and 28S): Conflict and agreement with the current system of classification", Molecular Phylogenetics and Evolution, 41 (2): 454–471, doi:10.1016/j.ympev.2006.05.017, PMID16815045
^ abGarrison, Nicole L.; Rodriguez, Juanita; Agnarsson, Ingi; Coddington, Jonathan A.; Griswold, Charles E.; Hamilton, Christopher A.; Hedin, Marshal; Kocot, Kevin M.; Ledford, Joel M. & Bond, Jason E. (2016), "Spider phylogenomics: untangling the Spider Tree of Life", PeerJ, 4: e1719, doi:10.7717/peerj.1719, PMC4768681, PMID26925338
^ abBond, Jason E.; Hendrixson, Brent E.; Hamilton, Chris A. & Hedin, Marshal (2012), "A Reconsideration of the Classification of the Spider Infraorder Mygalomorphae (Arachnida: Araneae) Based on Three Nuclear Genes and Morphology", PLOS ONE, 7 (6): e38753, doi:10.1371/journal.pone.0038753, PMC3378619, PMID22723885