Ochrophyte

Ochrophyte
Gusta šuma morskih algi sa podlogom
Gusta šuma morskih algi sa podlogom
Naučna klasifikacija e
Domen: Eukaryota
Kladus: Sar
Kladus: Stramenopiles
Kladus: Gyrista
Podtip: Ochrophytina
Cavalier-Smith, 1995[1]
Klase
Sinonimi

Ochrophytes su grupa uglavnom fotosintetskih stramenopila (heterokonta), smeštenih ili u razdeo Ochrophyta ili podrazdeo Ochrophytina.[5][6][7] Njihov plastid je poreklom od crvenih algi.[8][7]

Klasifikacija grupe se još razvija. Prvobitno, okrofiti su smatrani razdelom koji se naziva Ochrophyta. Neki autori su ga podelili na dva podrayreda, Phaeista Cavalier-Smith 1995 (koji obuhvata Hypogyristea i Chrysista u nekim klasifikacijama, ili Limnista i Marista u drugima) i Khakista Cavalier-Smith, 2000 (sadrži Bolidophyceae i silikatne alge).[9]

Filogenija

Kladogram ispod prikazuje evolutivne odnose između svih klasa okrofita. Zasnovan je na najnovijim filogenetskim analizama:[10][7][11][12]

Ochrophytina
Chrysista
"SI klada"
Fucistia

Xanthophyceaežutozelene alge
Allorge ex Fritsch 1935

Chrysoparadoxophyceae
Wetherbee et al. 2019

Phaeosacciophyceae
R.A.Andersen, L.Graf & H.S.Yoon 2020

Schizocladiophyceae
Henry, Okuda & Kawai 2003

Phaeophyceaesmeđe alge
Kjellman 1891

Aurophyceae
Cavalier-Smith 2013

"PX klada"
Raphidoistia

Raphidomonadea
Chadefaud ex Silva 1980

"SII klada"

Olisthodiscophyceae
Barcytė, Eikrem & M. Eliáš 2021

Limnistia

Chrysophyceaezlatne alge
Pascher 1914

Synchromophyceae
Horn et al. 2007

Eustigmatophyceae
Hibberd & Leedale 1971

Diatomista
Hypogyrista

Dictyochophyceae
Silva 1980

Pinguiophyceae
Kawachi et al. 2002

Khakista

Bolidophyceae
Guillou & Chretiennot-Dinet 1999

BacillariophyceaeSilikatne alge
Dangeard 1933

"SIII klada"

Verovatna sestrinska klada Ochrophyta/Ochrophytina je Actinophryidae,[13] prvobitno klasifikovana kao heliozoanski protisti.[14]

Reference

  1. ^ Cavalier-Smith T (1995). „Zooflagellate phylogeny and classification”. Cytology. 37 (11): 1010—1029. PMID 8868448. 
  2. ^ Cavalier-Smith, T. (1986). The kingdom Chromista, origin and systematics. In: Round, F.E. and Chapman, D.J. (eds.). Progress in Phycological Research. 4: 309–347.
  3. ^ Reviers, B. de. (2006). Biologia e Filogenia das Algas. Editora Artmed, Porto Alegre, p. 157.
  4. ^ Reviers, B. de. (2006). Biologia e Filogenia das Algas. Editora Artmed, Porto Alegre, pp. 15-16.
  5. ^ Thomas Cavalier-Smith; Ema E.-Y. Chao (2006). „Phylogeny and megasystematics of phagotrophic heterokonts (kingdom Chromista)”. Journal of Molecular Evolution. 62 (4): 388—420. Bibcode:2006JMolE..62..388C. PMID 16557340. S2CID 29567514. doi:10.1007/s00239-004-0353-8. 
  6. ^ Ingvild Riisberga; Russell J. S. Orr; Ragnhild Kluge; Kamran Shalchian-Tabrizi; Holly A. Bowers; Vishwanath Patil; Bente Edvardsen; Kjetill S. Jakobsen (2009). „Seven gene phylogeny of heterokonts”. Protist. 160 (2): 191—204. PMID 19213601. doi:10.1016/j.protis.2008.11.004. 
  7. ^ а б в Cavalier-Smith, Thomas (2018). „Kingdom Chromista and its eight phyla: a new synthesis emphasising periplastid protein targeting, cytoskeletal and periplastid evolution, and ancient divergences”. Protoplasma. 255 (1): 297—357. PMC 5756292Слободан приступ. PMID 28875267. doi:10.1007/s00709-017-1147-3Слободан приступ. 
  8. ^ Ševčíková T, Horák A, Klimeš V, et al. (2015). „Updating algal evolutionary relationships through plastid genome sequencing: did alveolate plastids emerge through endosymbiosis of an ochrophyte?”. Sci Rep. 5: 10134. Bibcode:2015NatSR...510134S. PMC 4603697Слободан приступ. PMID 26017773. doi:10.1038/srep10134Слободан приступ. 
  9. ^ „Heterokontophyta”. SHIGEN. National Institute of Genetics. Приступљено 18. 6. 2009. 
  10. ^ Derelle R, López-García P, Timpano H, Moreira D (новембар 2016). „A Phylogenomic Framework to Study the Diversity and Evolution of Stramenopiles (=Heterokonts)”. Mol Biol Evol. 33 (11): 2890—2898. PMC 5482393Слободан приступ. PMID 27512113. doi:10.1093/molbev/msw168Слободан приступ. 
  11. ^ Graf L, Yang EC, Han KY, Küpper FC, Benes KM, Oyadomari JK, Herbert RJH, Verbruggen H, Wetherbee R, Andersen RA, Yoon HS (децембар 2020). „Multigene Phylogeny, Morphological Observation and Re-examination of the Literature Lead to the Description of the Phaeosacciophyceae Classis Nova and Four New Species of the Heterokontophyta SI Clade”. Protist. 171 (6): 125781. PMID 33278705. S2CID 227315556. doi:10.1016/j.protis.2020.125781Слободан приступ. 
  12. ^ Dovilė Barcytė; Wenche Eikrem; Anette Engesmo; Sergio Seoane; Jens Wohlmann; Aleš Horák; Tatiana Yurchenko; Marek Eliáš (2. 3. 2021). „Olisthodiscus represents a new class of Ochrophyta”. Journal of Phycology. 57 (4): 1094—1118. PMID 33655496. doi:10.1111/jpy.13155Слободан приступ. hdl:10852/86515Слободан приступ. 
  13. ^ Azuma, Tomonori; Pánek, Tomáš; Tice, Alexander K.; Kayama, Motoki; Kobayashi, Mayumi; Miyashita, Hideaki; Suzaki, Toshinobu; Yabuki, Akinori; Brown, Matthew W.; Kamikawa, Ryoma (10. 4. 2022). „An Enigmatic Stramenopile Sheds Light on Early Evolution in Ochrophyta Plastid Organellogenesis”. Molecular Biology and Evolution. 39 (4). PMC 9004409Слободан приступ. PMID 35348760. doi:10.1093/molbev/msac065. 
  14. ^ Mikrjukov, Kirill A.; Patterson, David J. (2001). „Taxonomy and phylogeny of Heliozoa. III. Actinophryids” (PDF). Acta Protozoologica. 40: 3—25. 

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